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<title>UTas ePrints - Diverse fossil epacrids (Styphelioideae; Ericaceae) from early Pleistocene sediments at Stony Creek Basin, Victoria, Australia</title>
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<meta content="Jordan, Gregory J." name="eprints.creators_name" />
<meta content="Bromfield, Kate E." name="eprints.creators_name" />
<meta content="Sniderman, J.M. Kale" name="eprints.creators_name" />
<meta content="Crayn, Darren M." name="eprints.creators_name" />
<meta content="greg.jordan@utas.edu.au" name="eprints.creators_id" />
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<meta content="kalesniderman@gmail.com" name="eprints.creators_id" />
<meta content="Darren.Crayn@rbgsyd.nsw.gov.au" name="eprints.creators_id" />
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<meta content="2007-09-06" name="eprints.datestamp" />
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<meta content="Diverse fossil epacrids (Styphelioideae; Ericaceae) from
early Pleistocene sediments at Stony Creek Basin, Victoria, Australia" name="eprints.title" />
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<meta content="Epacrid, Styphelioideae, evolutionary radiation, Quaternary, fossils, Australia, sclerophylly, scleromorphy" name="eprints.keywords" />
<meta content="There is intense current interest in the radiation of the scleromorphic groups that dominate the Australian flora, but at present, only Proteaceae and Casuarinaceae have fossil records detailed enough to provide useful evidence of the timing of these radiations. This paper records a diverse assemblage of fossil leaves of another major scleromorphic group, the epacrids (subfamily Styphelioideae of Ericaceae, formerly known as Epacridaceae). The fossils are from Stony Creek Basin, in the western uplands of Victoria, Australia, and are of earliest Pleistocene age (circa 1.6 million years old). They include 19 forms sufficiently distinct to constitute different species. This diversity is considerably greater than the extant diversity of epacrids in the region. Published taphonomic data are used to argue that the actual diversity of the source vegetation of the fossil flora may have been significantly greater and comparable to the current local species richness of the centres of diversity. Ten of the fossil species are assigned to the largest extant tribe (Styphelieae), eight are assigned to Epacrideae and one is assigned to Cosmelieae. This evidence is used to argue that substantial radiation of the epacrids had occurred by the beginning of the Pleistocene." name="eprints.abstract" />
<meta content="2007" name="eprints.date" />
<meta content="published" name="eprints.date_type" />
<meta content="International Journal of Plant Sciences" name="eprints.publication" />
<meta content="168" name="eprints.volume" />
<meta content="UNSPECIFIED" name="eprints.thesis_type" />
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<meta content="1058-5893" name="eprints.issn" />
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<meta content="Albrecht DE 1996 Epacridaceae. Pages 464-509 in NG Walsh, TJ Entwistle, eds. Flora of Victoria. Vol. 4, Inkata Press, Melbourne, Australia.
Buchanan AM 2005 A census of the vascular plants of Tasmania and index to The Student's Flora of Tasmania. Tasmanian Herbarium Occasional Publication 7.
Carpenter RJ 1991 Palaeovegetation and Environment at Cethana, Tasmania. PhD diss. University of Tasmania, Hobart, Australia.
Carpenter RJ, Jordan GJ, Hill RS 1994 Banksieaephyllum taylorii (Proteaceae) from the Late Paleocene of New South Wales and its relevance to the origin of Australia's scleromorphic flora. Aust Syst Bot 7: 385-392.
Cherry W, Gadek PA, Brown EA, Heslewood MM, Quinn CJ 2001 Pentachondra dehiscens sp. nov. - an aberrant new member of Styphelieae. Aust Syst Bot 14: 513533.
Cranfield RJ 1998 Brachyloma nguba (Epacridaceae), a new species from the south-west of Western Australia. Nuytsia 12: 179183.
Cranfield RJ 2002 Conostephium magnum (Epacridaceae), a new species from Western Australia. Nuytsia 15: 2125.
Crayn DM, Kron KA, Gadek PA, Quinn CJ 1998 Phylogenetics and evolution of epacrids: a molecular analysis using the plastid gene rbcL with a reappraisal of the position of Lebetanthus. Aust J Bot 46: 187-200.
Crayn DM, Brown EA, Powell JM 2003 A revision of Lissanthe (Styphelioideae: Ericaceae). Aust Syst Bot 16: 595619.
Crayn DM, Hislop M, Heslewood MM 2005 Additions to Lissanthe (Styphelioideae: Ericaceae) in Western Australia: L. synandra sp. nov. and L. pleurandroides comb. nov. Aust Syst Bot 18: 1-7.
Crisp M, Cook L, Steane DA 2004 Radiation of the Australian flora: what can comparisons of molecular phylogenies across multiple taxa tell us about the evolution of diversity in present-day communities? Phil. Trans. R. Soc. Lond. B 359: 15511571
de Lange PJ, Sawyer JWD, Rolfe JR 2006 New Zealand Indigenous Vascular Plant Checklist. New Zealand Plant Conservation Network, Wellington, New Zealand. 94 pp.
Dettmann ME 1994 Cretaceous vegetation: the microfossil record. Pages 143-170 in RS Hill, ed. History of the Australian Vegetation: Cretaceous to Recent. Cambridge University Press, Cambridge.
Frakes LA 1999 Evolution of Australian environments. Flora of Australia 1 (2nd edn) 163-203.
Gradstein, FM, Ogg, JG, Smith, AG 2004 A Geological Time Scale 2004. Cambridge University Press, Cambridge.
Hill RS 1994 The history of selected Australian taxa. Pages 390-420 in RS Hill, ed. History of the Australian Vegetation: Cretaceous to Recent. Cambridge University Press, Cambridge.
Hill RS 1998 Fossil evidence for the onset of xeromorphy and scleromorphy in Australian Proteaceae. Aust Syst Bot 11: 391400
Hill RS, Gibson N 1986 Distribution of potential macrofossils in Lake Dobson, Tasmania. J Ecol 74: 373-84.
Hill RS, Orchard AE 1999 Composition and endemism of vascular plants. Pages 89-124 in JB Reid, RS Hill, MJ Brown, MJ Hovenden, eds. Vegetation of Tasmania. Australian Biological Resource Study, Melbourne, Australia.
Hopper SD, Gioia P 2004 The Southwest Australian floristic region: evolution and conservation of a global hot spot of biodiversity. Ann Rev Ecol Evol Syst 35: 623650.
Jackson WD 1999 The Tasmanian environment. Pages 1-10 in JB Reid, RS Hill, MJ Brown, MJ Hovenden, eds. Vegetation of Tasmania. Australian Biological Resource Study, Melbourne, Australia.
Jordan GJ 1997 Evidence of Pleistocene plant extinction and diversity from Regatta Point, western Tasmania, Australia. Bot J Linn Soc 123: 45-71.
Jordan GJ 2001 An investigation of long-distance dispersal based on species native to both Tasmania and New Zealand. Aust J Bot 49: 333-340.
Jordan GJ, Carpenter RJ, Hill RS 1998 Macrofossil evidence of past diversity of Proteaceae in Tasmania, including nine new species. Aust Syst Bot 11: 465-501.
Jordan GJ, Hill RS 1995 Oligocene leaves of Epacridaceae from Little Rapid River, Tasmania and the identification of fossil Epacridaceae leaves. Aust Syst Bot 8: 71-83.
Jordan GJ, Hill RS 1996 The fossil record of the Epacridaceae. Ann Bot 77: 341-346.
Keighery GJ 1996. Phytogeography, biology and conservation of Western Australian Epacridaceae. Ann Bot 77: 347-356.
Keighery GJ 2002 A new species of Conostephium (Epacridaceae) from gypsum dunes in Western Australia. Nordic J Bot 22: 4952.
Klak C, Reeves G, Hedderson T 2004. Unmatched tempo of evolution in Southern African semi-desert ice plants. Nature 427: 63-65.
Kron KA, Judd WS, Stevens PF, Crayn DM, Anderberg AA, Gadek PA, Quinn CJ, Luteyn JL 2002 Phylogenetic classification of Ericaceae: Molecular and morphological evidence. Bot Rev 68: 335423.
Lisiecki LE, Raymo ME 2005 A Pliocene-Pleistocene stack of 57 globally distributed benthic -18O records. Paleoceanography 20: 1-17.
Powell JM 1992 Epacridaceae. Pages 401-434 in GJ Harden, ed. Flora of NSW. University of NSW Press, Sydney, Australia
Quinn CJ, Brown EA, Heslewood MM, Crayn DM 2005 Generic concepts in Styphelieae (Ericaceae): the Cyathodes group. Aust Syst Bot 18: 439-454.
Scriven LJ, Hill RS 1995 Macrofossil Casuarinaceae: their identification, including the oldest macrofossil record, Gymnostoma antiquium sp. nov. from the Late Paleocene of New South Wales, Australia. Aust Syst Bot 8: 1035-1053.
Sniderman JMK, Pillans B, O'Suillivan PB, Kershaw AP 2007 Climate and vegetation in southeastern Australia respond to southern hemisphere insolation forcing in the Late Pliocene-Early Pleistocene. Geology 35:41-44.
Stevens PF, Luteyn J, Oliver EGH, Bell TL, Brown EA, Crowden RK, George AS, Jordan GJ, Ladd P, Lemson K, McLean CB, Menadue Y, Pate JS, Stace HM, Weiller CM 2004. Ericaceae. Pages 145-194 in K Kubitzki, ed. The Families and Genera of Vascular Plants. Vol. 6. Springer-Verlag, Berlin..
Taaffe G, Brown EA, Crayn DM, Gadek PA, Quinn CJ 2001 Generic concepts in Styphelieae: resolving the limits of Leucopogon. Aust J Bot 49: 107120.
Watson L 1967 Taxonomic implications of a comparative anatomical study of Epacridaceae. New Phyt 66: 495-504.
Weiller CM 1999 Leptecophylla, a new genus for species formerly included in Cyathodes (Epacridaceae). Muelleria 12: 195214." name="eprints.referencetext" />
<meta content="Jordan, Gregory J. and Bromfield, Kate E. and Sniderman, J.M. Kale and Crayn, Darren M. (2007) Diverse fossil epacrids (Styphelioideae; Ericaceae) from early Pleistocene sediments at Stony Creek Basin, Victoria, Australia. International Journal of Plant Sciences, 168 . ISSN 1058-5893 (In Press)" name="eprints.citation" />
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early Pleistocene sediments at Stony Creek Basin, Victoria, Australia" name="DC.title" />
<meta content="Jordan, Gregory J." name="DC.creator" />
<meta content="Bromfield, Kate E." name="DC.creator" />
<meta content="Sniderman, J.M. Kale" name="DC.creator" />
<meta content="Crayn, Darren M." name="DC.creator" />
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<meta content="There is intense current interest in the radiation of the scleromorphic groups that dominate the Australian flora, but at present, only Proteaceae and Casuarinaceae have fossil records detailed enough to provide useful evidence of the timing of these radiations. This paper records a diverse assemblage of fossil leaves of another major scleromorphic group, the epacrids (subfamily Styphelioideae of Ericaceae, formerly known as Epacridaceae). The fossils are from Stony Creek Basin, in the western uplands of Victoria, Australia, and are of earliest Pleistocene age (circa 1.6 million years old). They include 19 forms sufficiently distinct to constitute different species. This diversity is considerably greater than the extant diversity of epacrids in the region. Published taphonomic data are used to argue that the actual diversity of the source vegetation of the fossil flora may have been significantly greater and comparable to the current local species richness of the centres of diversity. Ten of the fossil species are assigned to the largest extant tribe (Styphelieae), eight are assigned to Epacrideae and one is assigned to Cosmelieae. This evidence is used to argue that substantial radiation of the epacrids had occurred by the beginning of the Pleistocene." name="DC.description" />
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<h1 class="ep_tm_pagetitle">Diverse fossil epacrids (Styphelioideae; Ericaceae) from early Pleistocene sediments at Stony Creek Basin, Victoria, Australia</h1>
<p style="margin-bottom: 1em" class="not_ep_block"><span class="person_name">Jordan, Gregory J.</span> and <span class="person_name">Bromfield, Kate E.</span> and <span class="person_name">Sniderman, J.M. Kale</span> and <span class="person_name">Crayn, Darren M.</span> (2007) <xhtml:em>Diverse fossil epacrids (Styphelioideae; Ericaceae) from early Pleistocene sediments at Stony Creek Basin, Victoria, Australia.</xhtml:em> International Journal of Plant Sciences, 168 . ISSN 1058-5893 (In Press)</p><p style="margin-bottom: 1em" class="not_ep_block"></p><table style="margin-bottom: 1em" class="not_ep_block"><tr><td valign="top" style="text-align:center"><a onmouseover="EPJS_ShowPreview( event, 'doc_preview_2238' );" href="http://eprints.utas.edu.au/1746/1/SCB_epacrids.pdf" onmouseout="EPJS_HidePreview( event, 'doc_preview_2238' );"><img alt="[img]" src="http://eprints.utas.edu.au/style/images/fileicons/application_pdf.png" class="ep_doc_icon" border="0" /></a><div class="ep_preview" id="doc_preview_2238"><table><tr><td><img alt="" src="http://eprints.utas.edu.au/1746/thumbnails/1/preview.png" class="ep_preview_image" border="0" /><div class="ep_preview_title">Preview</div></td></tr></table></div></td><td valign="top"><a href="http://eprints.utas.edu.au/1746/1/SCB_epacrids.pdf"><span class="ep_document_citation">PDF (Author Version - large file with high resolution images)</span></a> - Requires a PDF viewer<br />22Mb</td></tr><tr><td valign="top" style="text-align:center"><a onmouseover="EPJS_ShowPreview( event, 'doc_preview_2239' );" href="http://eprints.utas.edu.au/1746/2/SCB_Epacrid_preprint.pdf" onmouseout="EPJS_HidePreview( event, 'doc_preview_2239' );"><img alt="[img]" src="http://eprints.utas.edu.au/style/images/fileicons/application_pdf.png" class="ep_doc_icon" border="0" /></a><div class="ep_preview" id="doc_preview_2239"><table><tr><td><img alt="" src="http://eprints.utas.edu.au/1746/thumbnails/2/preview.png" class="ep_preview_image" border="0" /><div class="ep_preview_title">Preview</div></td></tr></table></div></td><td valign="top"><a href="http://eprints.utas.edu.au/1746/2/SCB_Epacrid_preprint.pdf"><span class="ep_document_citation">PDF (Author Version - smaller file with low resolution images)</span></a> - Requires a PDF viewer<br />5Mb</td></tr></table><p style="margin-bottom: 1em" class="not_ep_block">Official URL: <a href="http://www.journals.uchicago.edu/IJPS/home.html">http://www.journals.uchicago.edu/IJPS/home.html</a></p><div class="not_ep_block"><h2>Abstract</h2><p style="padding-bottom: 16px; text-align: left; margin: 1em auto 0em auto">There is intense current interest in the radiation of the scleromorphic groups that dominate the Australian flora, but at present, only Proteaceae and Casuarinaceae have fossil records detailed enough to provide useful evidence of the timing of these radiations. This paper records a diverse assemblage of fossil leaves of another major scleromorphic group, the epacrids (subfamily Styphelioideae of Ericaceae, formerly known as Epacridaceae). The fossils are from Stony Creek Basin, in the western uplands of Victoria, Australia, and are of earliest Pleistocene age (circa 1.6 million years old). They include 19 forms sufficiently distinct to constitute different species. This diversity is considerably greater than the extant diversity of epacrids in the region. Published taphonomic data are used to argue that the actual diversity of the source vegetation of the fossil flora may have been significantly greater and comparable to the current local species richness of the centres of diversity. Ten of the fossil species are assigned to the largest extant tribe (Styphelieae), eight are assigned to Epacrideae and one is assigned to Cosmelieae. This evidence is used to argue that substantial radiation of the epacrids had occurred by the beginning of the Pleistocene.</p></div><table style="margin-bottom: 1em" cellpadding="3" class="not_ep_block" border="0"><tr><th valign="top" class="ep_row">Item Type:</th><td valign="top" class="ep_row">Article</td></tr><tr><th valign="top" class="ep_row">Keywords:</th><td valign="top" class="ep_row">Epacrid, Styphelioideae, evolutionary radiation, Quaternary, fossils, Australia, sclerophylly, scleromorphy</td></tr><tr><th valign="top" class="ep_row">Subjects:</th><td valign="top" class="ep_row"><a href="http://eprints.utas.edu.au/view/subjects/260112.html">260000 Earth Sciences > 260100 Geology > 260112 Palaeontology</a><br /><a href="http://eprints.utas.edu.au/view/subjects/270401.html">270000 Biological Sciences > 270400 Botany > 270401 Plant Systematics, Taxonomy and Phylogeny</a></td></tr><tr><th valign="top" class="ep_row">ID Code:</th><td valign="top" class="ep_row">1746</td></tr><tr><th valign="top" class="ep_row">Deposited By:</th><td valign="top" class="ep_row"><span class="ep_name_citation"><span class="person_name">dr gregory j jordan</span></span></td></tr><tr><th valign="top" class="ep_row">Deposited On:</th><td valign="top" class="ep_row">06 Sep 2007</td></tr><tr><th valign="top" class="ep_row">Last Modified:</th><td valign="top" class="ep_row">09 Jan 2008 02:30</td></tr><tr><th valign="top" class="ep_row">ePrint Statistics:</th><td valign="top" class="ep_row"><a target="ePrintStats" href="/es/index.php?action=show_detail_eprint;id=1746;">View statistics for this ePrint</a></td></tr></table><p align="right">Repository Staff Only: <a href="http://eprints.utas.edu.au/cgi/users/home?screen=EPrint::View&eprintid=1746">item control page</a></p>
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